Artinskian (Early Permian) fusuline fauna from the Rongma area in northern Tibet: palaeoclimatic and palaeobiogeographic implications

A fusuline fauna consisting of ten species in five genera from the Qudi Formation of the Jiaomuri and Gangtangcuo sections in the Rongma area, northern Tibet, is described. This fauna contains a few typical Peri-Gondwanan fusuline species such as Pseudofusulina pamirensis, Neodutkevitchia insignis and N. sp. cf. N. tumidiscula. The fauna is dated as Artinskian based on the presence of both genera Chalaroschwagerina and Praeskinnerella. The Peri-Gondwanan fusuline fauna in the Qiangtang Block is grouped palaeobiogeographically into the Southern Transitional Zone, which is located in the southern part of the Peri-Gondwanan blocks during the late Sakmarian and Artinskian. Moreover, the occurrence of the Peri-Gondwanan fusuline fauna, the lithological transition from glacimarine deposits to carbonates, and the Peri-Gondwanan fusulines' apparent southeastward migration from the Kalmard Block of central Iran during the Artinskian, are interpreted here to be the result of global warming after the peak of the Late Palaeozoic Ice Age.